additions to Epler's 2010 "Water Beetles of Florida"
(last update 10 March
p. 13.11 - Pronoterus
has been synonymized with Suphisellus.
See Baca, S.M, Toussaint, E.F.A., Miller, K.B. & Short,
A.E.Z. 2017. Molecular phylogeny of the aquatic beetle
family Noteridae (Coleoptera: Adephaga) with an emphasis on data
partitioning strategies. Molecular Phylogenetics and
Evolution 107: 282-292. Thus our single species
of this former genus is now Suphisellus semipunctatus
pp. 6.6.15, 6.17 - Promoresia has been
synonymized with Optioservus.
See Kamite, Y. 2016. Revision
of the genus Optioservus Sanderson, 1953, part 3: The
O. elegans species group. Koleopterologische
Rundschau 86: 205-212. Thus we now have two species of Optioservus
recorded from Florida: O. ovalis (LeConte) and O.
pp. 11.1-11.9 - The species of
Hydrochus have been reviewed for
Mississippi (Worthington, R.J., Hellman, J.L. & Lago,
P.K. 2016. Hydrochidae (Coleoptera) of
Mississippi. Transactions of the American Entomological
Society 142: 167-213. As a result, several of my
letter-designated taxa now have "real" names. H.
sp 1 = H. falsus Hellman; H. sp. 2 = H.
jiawanae Makhan; H. sp. 5 = H. pajnii
Makhan; H. sp. 6 = H. schereri Makhan; H.
sp. 8 = H. jaechi Makhan.
pp. 5.61; 5.110 - Prodaticus has been reduced
to subgeneric status within Hydaticus.
Thus our most common species is now Hydaticus (Prodaticus)
bimarginatus. See Miller &
Bergsten. 2016. Diving Beetles of the
World. Systematics and Biology of the Dytiscidae. Johns
Hopkins University Press. 320 pp.
p. 5.73-5.74 - Coelambus Thomson has been reinstated as a full genus.
Thus two species previously placed in Hygrotus
are now in Coelambus: C. berneri (Young
& Wolfe) and C. nubilus (LeConte).
Note also that "nubilus" was misspelled in the
manual. See Miller, K.B. & Bergsten, J.
2016. Diving Beetles of the World. Systematics and
Biology of the Dytiscidae. Johns Hopkins University
Press. 320 pp.
p. 1.16 - There's a typo in the
first part of couplet 14 leading to Ptilodactylidae. The last
phrase should read "antennal segments 1 and 2 long, 3 very
short". Thanks to Doug Strom for informing me of my lack of
p. 5.78 - The genus Laccomimus
has been established by
Toledo & Michat. This genus replaces Laccodytes
Régimbart for the single species known from Florida, now known as Laccomimus
pumilio (LeConte). See Toledo, M. & Michat, M. 2015.
Description of Laccomimus gen. n. and eleven new species
from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae).
Zootaxa 3990: 301-354.
pp. 12.8; 12.16, 12.17 - Crenitulus
has been considered a synonym of Anacaena for some
time, but was recently reinstated as a full genus by Fikáček &
Vondráček (2014). Thus Anacaena
suturalis (LeConte) becomes Crenitulus suturalis
(LeConte). Anacaena limbata
remains the same, and is still unknown from Florida.
For more information and a corrected key separating the larvae
Crenitulus and Paracymus download
pp. 16.6-16.13 - Most species
from Florida are now placed in Contacyphon.
Zwick, P., Klausnitzer, B. & Ruta, R. 2013. Contacyphon Gozis, 1886 removed
from synonymy (Coleoptera: Scirtidae) to accommodate species so far
combined with the invalid name, Cyphon
Paykull, 1799. Entomologische Blätter und Coleoptera 109:
337-353. Note also that species from the "Cyphon collaris complex" (none known
to date from Florida) are now placed in the genus Nyholmia. See
Klausnitzer, B. 2013. Monophyletische Artengruppe der
ehemaligen Gattung Cyphon
Paykull, 1799 und Beschreibung von neuen Gattungen (Coleoptera,
Scirtidae). Entomologische Nachrichten und Berichte 57:
pp. 12.6, 12.30 - On p. 12.6
in the key for larval hydrophilids, the third statement in couplet
12 leading to Enochrus should read "LEFT mandible with one inner tooth,
RIGHT with two inner teeth". The figures are oriented
correctly. In the diagnosis for larvae on p. 12.30, the same
mistake was repeated. It is the right mandible that bears two
inner teeth; the left mandible bears one inner tooth. I wonder why
it took so long to find this boo-boo??
p. 16.52 - Ora discoidea Champion has been found
in Florida (Collier Co., Immokalee); it will key to O. texana but is marked differently.
It can be either mostly tan above or have mostly dark elytra
with a wide pale border, as in the Florida specimen here.
Thanks to Mike Thomas (FSCA) for the photo!
pp. 16.22-16.23 - Scirtes goliai Epler described, replaces Scirtes sp. 1. See Epler,
J.H. 2012. A new species of Scirtes (Coleoptera: Scirtidae) from southern
Florida and the Caribbean. Zootaxa 3530: 77-82.
pp. 5.38, 18.2, 18.7 -
parentheses removed from author name for Celina grossula - should be Celina grossula LeConte. I had
followed an error in Young (1979) by placing "LeConte" in
parentheses. Because LeConte originally described the species grossula in the genus Celina, there are no
parentheses around his name.
pp. 3.1, 3.7 - To reiterate
about the spelling of the name of the weevil Auleutes, it's Auleutes,
not Aleutes or Auletes.
p. 5.38, 5.39, 5.41 - Celina
slossonae changed back to its original spelling - call it
Celina slossoni ...
again. Note that even though Mutchler misspelled it as "slossoni" (which is
incorrect; because the species was named for a woman, it should
have been "slossonae"),
Article 33.4 of the ICZN states that such a change is to be termed
an "incorrect subsequent spelling".
p. 5.92 - the captions for the
two beetles are switched; N. p.
pullus is on the left, N.
p. floridanus on the right! Thanks to Dana Denson
(RCID) for catching this!
p. 5.26 - The
genus Stictotarsus is not known from
Florida. Because it is recorded from South Carolina and there
is thus a possibility it may occur here, I included the genus,
represented by the widespread taxon S. griseostriatus (De Geer), in my key to
genera of dytiscid adults (p. 5.26) of Florida.
It has long been known that the taxon is notable for its variation
and is most likely a complex of species, especially in Europe. Now
this taxon has been established as a new genus, Boreonectes (Angus, R.B. 2010. Boreonectes gen. n., a new
genus for the Stictotarsus
griseostriatus (De Geer) group of sibling species
(Coleoptera: Dytiscidae), with additional karyosystematic data on
the group. Comparative Cytogenetics 4: 123-131). Thus, the
species’ name is now Boreonectes griseostriatus (De
Geer). Given the variability of this taxon, it is most likely
that at least one of the “variants” known from North America
actually represents an undescribed species, so stay tuned.
Also note that some other Nearctic Stictotarsus are now included in Boreonectes. These are: B. aequinoctialis (Clark), B. coelamboides (Fall), B. dolerosus (Leech), B. expositus (Fall), B. funereus (Crotch), B. panaminti (Fall) (tentative
placement), B. spenceri (Leech)
B. striatellus (LeConte).
Boreonectes may be
separated from Stictotarsus
by the shining ventral surface of the head behind the eyes,
with only superficial reticulation (closely punctate in Stictotarsus).
p. 1.6 - 2nd column,
under 3., line 15 should read “the bulging, white abdominal venter,
p. 1.8 - Brady Richards (California State University, Chico, CA) was
somehow left off the list of workers who supplied specimens – sorry,
pp. 3.1, 3.7 – Auleutes is
misspelled as “Aleutes”.
p. 5.63 - Couplet 1 has H.
deflatus at 3.8 – 4.7mm but the description has it listed
as 4.2-4.7mm; go with 3.8-4.7 mm.
p. 5.70 - species box: delete space in “inexpectatus”
p. 6.23 – what are termed “basolateral” triangles in couplet 14
should be basomesal triangles
p. 16.14 - under Notes – the “fourth form” referred to in line
1 is not “keyed below”; it should read “discussed below”.