p 3.21 - Agabus has been split into Agabus and Platambus. For adults, the two genera will separate at the first couplet, with Agabus going to couplet 2 and Platambus going to couplet 3. For larvae, they may separate at the first couplet – Agabus species would key to couplet two; Platambus species to couplet one. Agabus aeruginosus Aubé has been split. The species we have in Florida is now called Agabus xyztrus Larson. The undescribed species referred to in the manual as Agabus sp. A, described as A. astrictovittatus Larson & Wolfe in Larson & Wolfe (1998), is now named Platambus astrictovittatus (Larson & Wolfe).
p 3.23 - The species Agaporomorphus dodgei Young has been moved to a new genus and is now called Hydrodytes dodgei (Young). See Miller (2001).
pp 3.36-3.37 - Coptotomus lenticus Hilsenhoff is considered a subspecies of C. longulus LeConte by Larson et al. (2000); thus the species/subspecies in Florida is C. longulus lenticus Hilsenhoff.
pp 3.56-3.57 - The Hydroporus oblitus group is established as a new genus, Hydrocolus Larson & Roughley.
pp 3.70-3.72 - Liodessus affinis is a complex of
species; the species we have in Florida is Liodessus noviaffinis
Miller. Liodessus fuscatus
has been renamed L. crotchi
p 3.76 - Couplet 1 of key should read: "Length less than 6.5 mm;" Under Notes on species, length for M. leechi should be 5.5-6.1 mm.
p 3.81 - Couplet 7 should read: "Length < 4.1 mm;"
p 3.83 - Couplet 13' should read: "Length larger, 3.1-4.6 mm;"
p 3.84 - Couplet 22 should read: "Smaller, length 3.1-3.8 mm;"
p 3.85 - Under Notes on species, length for N. asidytus should be 3.1-3.8 mm.
p 3.86 - Length for N. hybridus should read: "around 3.5 mm"
p 3.90 - The characters given for the separation of the
larvae of Thermonectus basillaris and Th. ornaticollis will
work. Wilson’s (1923) original description of the larvae was
At this time larvae may be identified only to genus.
Larvae and adults of Oulimnius latiusculus (LeConte) have
in western Florida. In my elmid key the adult will key to couplet 4, Microcylloepus.
Oulimnius lacks the serrated lateral margins of the pronotum
on Microcylloepus. Oulimnius is also more oval in shape
the tooth that clasps the epipleuron arises from the margin of the 4th
sternite; in Microcylloepus this tooth arises from the 5th
Larvae of Oulimnius will key to couplet 7; they can be
Macronychus by the pleurae on abdominal segments 1-7.
are separated from Gonielmis by the lack of dorsal,
humps on the abdominal segments and by having the anterior and
portions of the mesopleural sclerites subequal - in Gonielmis
mesopleural sclerite is much smaller and thinner than the posterior
A second species of Oulimnius,
O. nitidulus (LeConte) has
been collected in the Panhandle from a stream on Eglin AFB. The
two species are easy to separate:
O. latiusculus - Length
1.50 mm; elytra with series of regular striae with equally sized pits.
O. nitidulus - Length 1.25 mm; elytra with irregular striae, with some pits much larger than others.
p 8.27 - The correct name for Enochrus perplexus
is Enochrus fimbriatus (Melsheimer). Although Gundersen
the synonymy of the two names, he chose the junior synonym and thus the
name perplexus (LeConte).
8(7’) Ligula shorter than
or subequal to basal segment of labial palp .. 8a
8’ Ligula longer than basal segment of labial palp ………………. 10
8a(8) Right mandible with 2
inner teeth; left mandible with one inner tooth .. Enochrus
8a’ Both mandibles with 2 inner teeth ……………………………. 9
p 8.9 - A caution for users: Couplet 11' should read "Mid and hind tibiae without a fringe of swimming setae (although setae may be present on mid and hind tarsi only)". Users may mistakenly key Hydrobius to couplet 12 because Hydrobius tumidus has fine natatory setae on its mid and hind tarsi but none on the tibiae. Of course, Hydrobius will not fit the diagnosis for Berosus or Derallus!!
p 8.10 - Couplet 13 should read “Maxillary palp short and stout, no more than length of antenna, with last segment at least as long as the preceding segment”
p 8.11 - Couplet 17' should read "Mid and hind tarsi
p 8.16 - Berosus striatus (Say) is now called B.
p 8.41 - Makhan (2001) recently established the
Rishihydroius, which includes several species formerly placed in
As far the SE US fauna is concerned, he included in the new genus Hydrochus
callosus LeConte, which would become R. callosus (LeConte).
taxa included are R. brevitarsus (Knisch), R. currani
R. scabratus (Mulsant) and R. setosus (Leach). The taxa Hydrochus
sp. 1 and H. sp. 2 from my manual would also be included in Rishihydroius.
HOWEVER - NOTE THAT NO HYDROPHILID WORKERS HAVE ACCEPTED MAKHAN’S NAMES
OR CLASSIFICATIONS!! Also note that the Hydrochidae is now
considered a separate family, as are the Helophoridae.
Musgrave, P.N. 1935. A synopsis of the genus Helichus
Erichson in the United States and Canada, with description of a new
species (Coleoptera: Dryopidae). Proc. Ent. Soc. Wash. 37:
Archangelsky, M. 1997. Studies on the biology, ecology and systematics of the immature stages of New World Hydrophiloidea (Coleoptera: Staphyliniformia). Ohio Biological Survey Bulletin New Series Vol. 12 No. 1. ix + 207pp.
Arnett, R.H., Jr. & M.C. Thomas (eds.). 2001. American Beetles. Volume 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press. Boca Raton, London, New York, Washington, D.C. 443 pp.
Bacon, M.A., E.H. Barman & B.P. White. 2000. Biology of Coptotomus lenticus (Coleoptera: Dytiscidae: Coptotominae) with a description of its mature larva. J. Elisha Mitchell Sci. Soc. 116: 75-81.
Barman, E.H., M.E. Blair & M.A. Bacon. 2001. Biology of Ilybius
oblitus (Coleoptera: Dytiscidae) with a description of its mature
larva and an evaluation of diagnostic characters for separation of
southeastern Ilybius and Agabus larvae. J. Elisha
Mitchell Sci. Soc. 117: 81-89.
Ciegler, J.C. 2003. Water Beetles of South Carolina
(Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae,
Hydrophilidae, Hydraenidae, Scirtidae, Elmidae, Dryopidae, Limnichidae,
Heteroceridae, Psephenidae, Ptilodactylidae, and Chelonariidae). Biota
of South Carolina, Vol. 3. Clemson University, Clemson, S.C. 207
pp. Costs $40.00 (incl. S/H). It may be ordered online: http://cufan.clemson.edu/olos/
Logon as GUEST and search the term "Biota".
Or send check payable to Clemson University to: Public Service Bulletin Room, 96 Poole Agricultural Center, Clemson University, Clemson, SC 29634-0129
Hansen, M. 1999. Hydrophilodea (s. str.) (Coleoptera). In World Catalogue of Insects, Vol.
2: 1-416. Apollo Books, Stenstrup.
Larson, D.J., Alarie, Y. and Roughley, R.E. 2000. Predaceous Diving Beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska. National Research Council of Canada Research Press, Ottawa, Ontario, Canada, 982 pp. ISBN 0-660-17967-9. US $64.95. For ordering info, go to: http://www.monographs.nrc.ca
Larson, D.J. & G.W. Wolfe. 1998. Revision of the North American Agabus (Coleoptera: Dytiscidae): the semivittatus group. Can. Ent. 130: 27-54.
Makhan, D. 2001. A new genus Rishihydroius and three new Hydrochus species (Coleoptera: Hydrochidae) from Africa. Bangladesh J. Zool. 29: 83-95.
Miller, K.B. 1998. Revision of the Nearctic Liodessus affinis (Say 1823) species group (Coleoptera: Dytiscidae, Hydroporinae, Bidessini). Ent. scand. 29: 281-314.
Miller, K.B. 2001. On the phylogeny of the Dytiscidae (Insecta:
Coleoptera) with emphasis on the morphology of the female reproductive
tract. Insect Syst. Evol. 32: 45-92
Nilsson, A.N. 2000. A new view on the generic classification of the Agabus-group of genera of the
Agabini, aimed at solving the problem with a paraphyletic Agabus. Koleopterische Rundschau
Nilsson, A.N. 2001. Dytiscidae. In
World Catalogue of Insects, Vol. 3: 1-395. Apollo Books, Stenstrup.
Larson, D.J., Alarie, Y. and Roughley, R.E. 2000. Predaceous
Diving Beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with
emphasis on the fauna of Canada and Alaska. National Research Council of Canada Research Press, Ottawa, Ontario, Canada, 982 pp.
64.95$ CAN, other countries 64.95$ US.
For ordering info, go to:
This publication is a MASTERPIECE! Anybody seriously interested in Dytiscidae must obtain it!!
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